PANNARIA Delise
Pannaria conoplea - cited from Missouri by the following sources: Berry (1937), from Franklin, Jefferson, and St. Louis cos. [as P. rubiginosa var. lanuginosa]; Gier & Kendrick (1972), from Jefferson Co. and also a literature report [as P. pityrea]; Hale (1969), mapped from extreme SE Mo. [as P. pityrea] - note that Hale (1979) maps this species from no closer to Missouri than the central Appalachians; and Nelson (1911), from Jefferson and St. Louis cos. [as P. lanuginosa]. Considerable confusion surrounds these reports, and no confirming specimens have been located, nor is this species known from any states bordering Missouri. Gier & Kendrick (1972) refer the report of Crocynia lanuginosa in Gier & Jones (1954) to this species [as P. pityrea], but the name is actually a synonym of Lepraria membranacea; see discussion under that species. To further complicate matters, a specimen from St. Louis Co. labelled P. lanuginosa, Russell 18 (MICH), is Lepraria lobificans! Based on the apparent confusion with Lepraria that has surrounded this taxon in Missouri, the Berry report may also be referable to Lepraria; his habitat description of "sheltered limestone ledges and bases of trees" would seem to support this.
Pannaria fallax [sensu Gier & Jones (1954) and Gier & Kendrick (1972)] see Arthopyrenia lapponina
Pannaria lanuginosa
Pannaria conoplea
Pannaria leucosticta
Fuscopannaria leucosticta
Pannaria lurida (Mont.) Nyl.
Berry (1937): Franklin Co.
C. Culberson (1969): literature report.
Fink (1935).
Gier & Kendrick (1972): Barry Co., also literature report.
Hale (1957a): Barry, Dent, Douglas, Ozark, and Pulaski cos.
Hale (1969, 1979): mapped from extreme S Mo.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Mohr (1901) [as Physma luridum].
Wetmore (1992a): Taney Co.
Pannaria microphylla
Fuscopannaria leucophaea
Pannaria pityrea
Pannaria conoplea
Pannaria rubiginosa (Ach.) Bory
Gier & Kendrick (1972): literature report.
Hale (1969): mapped from all but N1/4 Mo.; note that Hale's 1979 revision maps the species from no closer to Missouri than the Appalachians.
Ladd (1991): Ripley Co.; designated as Endangered.
MDC (1991, 1992): designated as Endangered.
Nigh et al. (1992): Ripley Co.
Pannaria rubiginosa var. lanuginosa
Pannaria conoplea
Pannaria tavaresii - although mapped from southern Missouri in the shaded range map in Hale (1979), there is no other information attributing this taxon to Missouri.
PARMELIA Ach.
Parmelia sp.
Redfearn et al. (1969): Texas Co.; this probably is referable to a Flavoparmelia or Xanthoparmelia species.
Thomas & Jackson (1985).
Parmelia acctabulum [
Pleurosticta accetabulum (Neck.) Elix & Lumbsch] - cited from Pike Co. by Pech (1866). According to Egan (1987), this lichen does not occur in North America.
Parmelia angustiphylla
Xanthoparmelia angustiphylla
Parmelia arnoldii [¸ Parmotrema arnoldii (Du Rietz) Hale] - cited from Camden Co. by Gier & Jones (1954) and Gier & Kendrick (1972). The specimen on which these reports are based, Jones 5158 (MCJ), is Parmotrema eurysacum.
Parmelia aspidota - see Parmelia exasperata
Parmelia aurulenta
Myelochroa aurulenta
Parmelia austrosinensis
Parmotrema austrosinense
Parmelia bolliana
Punctelia bolliana; note that early reports may include Punctelia semansiana. As used by W. Culberson & C. Culberson (1956), Parmelia bolliana strain 1 (lecanoric acid)
Punctelia semansiana and Parmelia bolliana strain 4 (protolichenesteric acid)
Punctelia bolliana.
Parmelia borreri [
Punctelia borreri (Sm.) Krog] - local reports of this species are referable to Punctelia subrudecta and/or Punctelia missouriensis. Berry (1936) describes this taxon as sorediate, suggesting his specimens may be referable to Punctelia missouriensis, but several Missouri collections by Berry determined as this species are Punctelia bolliana. Gier & Kendrick (1972) mention that some of the reports of this taxon by Gier & Jones (1954) are referable to Punctelia rudecta.
Parmelia candelaria - cited from Pike Co. by Pech (1866); this probably refers to a taxon of Candelaria or Xanthoria.
Parmelia caperata
Flavoparmelia caperata; most Missouri reports for Parmelia caperata include the morphology now separated as Flavoparmelia baltimorensis.
Parmelia capitata [sensu Hus (1908)]
Flavoparmelia caperata
Parmelia caroliniana
Canoparmelia caroliniana
Parmelia cetrata
Rimelia cetrata
Parmelia colpodes
Anzia colpodes
Parmelia conspersa (in part)
Xanthoparmelia conspersa; note that early reports of Parmelia conspersa are impossible to translate into modern taxonomic concepts, inasmuch as the name has been applied indiscriminately and inconsistently to a broad spectrum of Xanthoparmelia species. The issue becomes even more confused since many early reports apparently followed the delineations used by Hale (1955b), who considered Parmelia conspersa to refer to what are now segregated as various non-isidiate Xanthoparmelia species, including X. angustiphylla, X. cumberlandia, X. hypomelaena, X. somloënsis, and X. tasmanica. Lower thallus coloration, now considered an important taxonomic character, is not mentioned in most early treatments. Parmelia conspersa, stictic acid strain sensu Hale (1955b)
Xanthoparmelia angustiphylla and/or X. cumberlandia; Parmelia conspersa, salazinic acid strain sensu Hale (1955b)
Xanthoparmelia somloënsis and/or X. tasmanica; Parmelia conspersa, fumarprotocetraric acid strain sensu Hale (1955b)
Xanthoparmelia hypomelaena. As used by Berry (1941), this apparently refers to X. cumberlandia and/or X. somloënsis.
Parmelia convexiuscula - cited from Pike Co. by Pech (1866). This is apparently the nomenclatural equivalent of Anzia colpodes; Tuckerman (1882) lists Imbricaria convexiuscula as a synonym of Parmelia colpodes [¸ Anzia colpodes]. Given the uncertainty regarding many early lichen determinations and the lack of any Pech specimens from which to assess his taxonomic competence, this report is provisionally excluded. Modern data restrict A. colpodes to the southern portions of the state.
Parmelia crinita
Parmotrema crinitum
Parmelia cumberlandia
Xanthoparmelia cumberlandia
Parmelia cycloselis - cited from Pike Co. by Pech (1866); this is an apparent synonym of Phaeophyscia orbicularis (see Moberg 1977). As pointed out under Phaeophyscia orbicularis, early concepts in this group encompassed a complex of several species. Since the Pech specimens were destroyed, it is impossible to assign this report to any modern species concept.
Parmelia dilatata - local reports are referable to Parmotrema gardneri.
Parmelia dissecta
Parmelinopsis minarum
Parmelia erecta
Parmotrema perforatum; some early reports also include conceptually Parmotrema hypotropum - several early Missouri collections labelled as this taxon are Parmotrema hypotropum.
Parmelia eurysaca
Parmotrema eurysacum
Parmelia exasperata [
Melanelia exasperata (De Not.) Essl.] - cited from Missouri by no less than 5 references, but the bases for all of these reports are questionable. The Boone Co. report by Berry (1936) [as Parmelia aspidota] was not included in Berry (1937). Only one of the three specimens cited by Berry (1936) has been located, Berry 308 (UMO), which is a Collema. His description of this as a terricolous lichen with an olivaceous thallus suggests that all of his collections may be referable to Collema. Esslinger (1977b) includes possibly up to 3 mapped records from Missouri, but the exact locality and state are uncertain, and no Missouri specimens are cited. Gier & Kendrick (1972) include a literature report based on Hale (1969), whose range map includes SE Missouri. Subsequently, Hale (1979) changed his range map to include all but NW Mo. Since no confirming specimens have been located, the species is provisionally excluded from the flora.
Parmelia finkii
Myelochroa obsessa
Parmelia formosana - see Hypotrachyna formosana
Parmelia frondifera
Punctelia bolliana - local reports may include material referable to Punctelia semansiana.
Parmelia galbina
Myelochroa galbina
Parmelia horrescens
Parmelinopsis horrescens
Parmelia hyperleucites [sensu MBG (1974a, 1974b)]
Punctelia semansiana
Parmelia hypoleucites [
Punctelia hypoleucites (Nyl.) Krog] local reports are referable to Punctelia semansiana.
Parmelia hypomelaena
Xanthoparmelia hypomelaena
Parmelia hypopsila
Xanthoparmelia angustiphylla for North American reports.
Parmelia hypotropa
Parmotrema hypotropum
Parmelia hypotropoides
Parmotrema perforatum
Parmelia isidiata (in part - fumarprotocetraric acid strain) - local reports are referable to Xanthoparmelia subramigera.
Parmelia isidiosa (stictic acid strain) [sensu Hale (1955b)]
Xanthoparmelia conspersa and/or X. plittii
Parmelia laevigata [¸ Hypotrachyna laevigata (Sm.) Hale] - cited from Pulaski Co. by Berry (1941); this forms the basis for the literature reports by Gier & Jones (1954) and Gier & Kendrick (1972). Hale (1979) implies that this species is narrowly restricted to the southern Appalachians. Although the specimen has not been located, it is almost certain that the Missouri reports are based on misidentifications.
Parmelia leucochlora [
Pseudoparmelia sphaerospora (Nyl.) Hale] - cited from Missouri by Fink (1935). Hale (1979) maps this coastal plain species from Texas to southern North Carolina. Given the remoteness of the entire range of the species from Missouri and the lack of confirming specimens, this species is excluded from the flora.
Parmelia livida
Hypotrachyna livida
Parmelia louisianae
Parmotrema louisianae
Parmelia margaritata
Parmotrema margaritatum
Parmelia mexicana
Xanthoparmelia mexicana
Parmelia michauxiana
Parmotrema michauxianum
Parmelia obsessa
Myelochroa obsessa
Parmelia olivacea [
Melanelia olivacea (L.) Essl.] - cited from Franklin Co. by Berry (1937); Gier & Kendrick (1972) consider this record "doubtful". Esslinger (1977b) maps this species from no closer to Missouri than central Wisconsin, and no confirming specimens have been located, so the plant is excluded from the flora.
Parmelia perforata
Parmotrema perforatum; note that Hale (1957a) uses this name for material containing salazinic acid - such reports are referable to Parmotrema eurysacum or possibly Rimelia cetrata.
Parmelia perlata [
Parmotrema chinense (Osbeck) Hale & Ahti] - cited from Greene and St. Francois cos. by Berry (1941); this forms the basis for the literature report in Gier & Jones (1954), who also cite a specimen from Camden Co. (as Parmelia trichotera). Gier & Kendrick (1972) erroneously attribute a literature report to Fink (1907b). To date, all local material originally identified as this species is Parmotrema hypotropum or Rimelia reticulata. Hale (1979) maps this species in North America only from the Great Lakes, Appalachians, and Pacific coast.
Parmelia perreticulata
Punctelia perreticulata
Parmelia pertusa [
Menegazzia terebrata (Hoffm.) A. Massal.] - cited from Boone Co. by Berry (1936, 1937); Gier & Kendrick consider the 1937 reports "doubtful". Only one of the 3 specimens reported by Berry (1937) has been located. This collection, Berry 289 (UMO), is Flavoparmelia caperata. Since M. terebrata is a northern species whose range is remote from Missouri, it is excluded from the flora.
Parmelia physodes [
Hypogymnia physodes (L.) Nyl.] cited from Boone Co. by Berry (1936, 1937); Gier & Kendrick consider the 1937 report "doubtful". The only Berry determination of this taxon located, Berry 102 (UMO), is Physcia stellaris. Since the range of H. physodes is north of Missouri, it is excluded from the flora.
Parmelia plittii - see Xanthoparmelia plittii
Parmelia praesorediosa [
Parmotrema praesorediosum (Nyl.) Hale] cited from Vernon Co. by Gier & Kendrick (1972), who also erroneously attribute a literature report of this species to Hale (1969). The Vernon Co. report is apparently based on Kendrick 10984 (MCJ), which is Rimelia reticulata.
Parmelia quercina - Hale (1979) states that reports of this taxon from the eastern United States are referable to either Hypotrachyna livida or Myelochroa galbina.
Parmelia reticulata
Rimelia reticulata
Parmelia rudecta
Punctelia rudecta
Parmelia sacatilis - cited from Boone Co. by Berry (1936), apparently in reference to P. saxatilis. The specimen this report is based on, Berry 300 (UMO), is Xanthoparmelia angustiphylla.
Parmelia saxatilis - local reports of this species are referable to Parmelia squarrosa, although some Berry determinations as P. saxatilis are Punctelia rudecta.
Parmelia semansiana
Punctelia semansiana
Parmelia squarrosa Hale
Berry (1937): Boone, Callaway, and Franklin cos. [as P. saxatilis].
Berry (1941): Wayne Co. [as P. saxatilis].
Gier & Kendrick (1972): literature report [as P. saxatilis].
Hale (1979): mapped from S3/5 Mo.
Hale (1987): one specimen cited from "Missouri".
Ladd (1991): Dent, Madison, and Texas cos.; designated as Watch List.
MDC (1991, 1992): designated as Watch List.
Parmelia stenophylla (stictic acid strain) [sensu Hale (1955b)]
Xanthoparmelia angustiphylla and/or X. cumberlandia; Parmelia stenophylla (salazinic acid strain) [sensu Hale (1955b)]
Xanthoparmelia somloënsis and/or X. tasmanica.
Parmelia subcrinita - reported from Missouri by Hale (1957a, 1969); the 1957 Hale report is the basis for the literature report in Gier & Kendrick (1972). This name was used for species containing atranorin and salazinic acid. Unless the chemistry is fully explicated, there is no means of reliably translating old reports into modern nomenclature, since the reports could refer to Parmotrema ultralucens, Rimelia subisidiosa, or Rimeliella subtinctoria.
Parmelia sublaevigata
Hypotrachyna livida and/or Myelochroa galbina
Parmelia subquercifolia
Myelochroa galbina (probably including Hypotrachyna livida for local reports).
Parmelia subramigera
Xanthoparmelia subramigera
Parmelia subrudecta
Punctelia subrudecta
Parmelia subrugata - although cited from Boone Co. by Berry (1936), and from Boone, Johnson, and Washington cos. by Berry (1937), Gier & Kendrick (1972) consider these reports "doubtful". Esslinger & Egan (1995) consider reports of this species from North America to be misidentifications. The one Berry determination of this taxon located, Berry 397 (UMO), is Rimeliella subtinctoria.
Parmelia subtinctoria
Rimeliella subtinctoria
Parmelia sulcata Taylor
Hale (1979): mapped from SE1/4 Mo.
Ladd et al. (1994): Boone Co.
Parmelia sulphurosa
Myelochroa galbina
Parmelia taractica [
Xanthoparmelia taractica (Kremp.) Hale] - local reports are referable to Xanthoparmelia somloënsis. See note under Xanthoparmelia taractica.
Parmelia tasmanica
Xanthoparmelia tasmanica
Parmelia texana
Canoparmelia texana
Parmelia tiliacea
Hypotrachyna livida (possibly including Myelochroa galbina for local reports).
Parmelia tinctorum
Parmotrema tinctorum
Parmelia trichotera see Parmelia perlata
Parmelia ulotrix
Phaeophyscia ciliata
Parmelia xanthina - see Parmotrema xanthinum
Parmelina aurulenta
Myelochroa aurulenta
Parmelina dissecta
Parmelinopsis minarum
Parmelina galbina
Myelochroa galbina
Parmelina horrescens
Parmelinopsis horrescens
Parmelina obsessa
Myelochroa obsessa
Parmelinopsis horrescens (Taylor) Elix & Hale
Gier & Kendrick (1972): Jasper Co., also literature report [as Parmelia horrescens].
Hale (1969): mapped from SE2/3 Mo. [as Parmelia horrescens].
Hale (1979): mapped from all but NW Mo. [as Parmelina horrescens].
Ladd (1991): Carter Co.; designated as Endangered.
MDC (1991, 1992): designated as Endangered.
Parmelinopsis minarum (Vain.) Elix & Hale
Dey (1976, 1979): mapped from SE Mo., based on literature report [as Parmelia dissecta].
Gier & Kendrick (1972): Vernon Co., also literature report [as Parmelia dissecta].
Hale (1969): mapped from SE Mo. [as Parmelia dissecta].
Hale (1979): mapped from SE Mo. [as Parmelina dissecta].
PARMOTREMA A. Massal.
Parmotrema sp. [sensu Wilhelm & Ladd (1986)]
Parmotrema gardneri
Parmotrema austrosinense (Zahlbr.) Hale
C. Culberson (1969): literature report [as Parmelia austrosinensis].
Gier & Kendrick (1972): Newton Co., also literature report [as Parmelia austrosinensis].
Hale (1959): Taney Co. [as Parmelia austrosinensis].
Hale (1961) [as Parmelia austrosinensis].
Hale (1965): Taney Co. [as Parmelia austrosinensis].
Hale (1979): mapped from S1/3 Mo.
Skorepa (1973) [as Parmelia austrosinensis].
Parmotrema cetratum
Rimelia cetrata
Parmotrema crinitum (Ach.) M. Choisy
Berry (1941): Dunklin and Wright cos. [as Parmelia crinita].
C. Culberson (1969): literature report [as Parmelia crinita].
Gier & Jones (1954): literature report [as Parmelia crinita].
Gier & Kendrick (1972): literature report [as Parmelia crinita].
Hale (1957a): Douglas and Pulaski cos. [as Parmelia crinita].
Hale (1965): Douglas and Pulaski cos. [as Parmelia crinita].
Hale (1969): mapped from SE1/2 Mo. [as Parmelia crinita].
Hale (1979): mapped from SE Mo.
Ladd (1996): Phelps Co.
Wetmore (1992a): Taney Co. [as Parmelia crinita].
Parmotrema eurysacum (Hue) Hale
W. Culberson & Hale (1974): 3 records mapped from SW Mo. [as Parmelia eurysaca]
Hale (1979): mapped statewide.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Wetmore (1992a): Taney Co. [as Parmelia eurysaca]. Note: Hale (1957a) reports Parmelia perforata from Barry, Dent, Pulaski, Taney, and Wright cos.; these are listed as having salazinic acid, and are probably referable here, or possibly to Rimelia cetrata. See Rimelia cetrata and Parmelia arnoldii.
Parmotrema gardneri (C. W. Dodge) Sérus.
Gier & Kendrick (1972): Jasper Co. [as Parmelia dilatata].
Ladd & Wilhelm (1986): Barry, Butler, Lincoln, Shannon, and Wayne cos. [as Parmotrema sp.].
Wilhelm & Parker (1989): "Missouri Ozarks" [as Parmotrema submichauxianum]. See the discussion of this taxon in Harris (1990)].
Parmotrema haitiense
Rimeliella subtinctoria
Parmotrema hypoleucinum (J. Steiner) Hale
Ladd (1991): Osage Co.; designated as Status Undetermined.
Ladd & Wilhelm (1986): Osage Co.
MDC (1991, 1992): designated as Status Undetermined.
Parmotrema hypoleucites [sensu AES (1986)]
Punctelia semansiana
Parmotrema hypotropum (Nyl.) Hale
Berry (1936): Boone Co. [as Parmelia hypotropa].
Berry (1937): Boone Co. [as Parmelia hypotropa].
W. Culberson (1973): mapped from SE1/5 Mo. [as Parmelia hypotropa].
W. Culberson (1986): 4 map records SE Mo.
W. Culberson & C. Culberson (1973): 4 mapped records S Mo. [as Parmelia hypotropa].
Gier & Kendrick (1972): Newton Co., also literature report [as Parmelia hypotropa].
Hale (1965): Texas Co., also 4 mapped records SE1/2 Mo. [as Parmelia hypotropa].
Hale (1969): mapped from SE1/2 Mo. [as Parmelia hypotropa].
Hale (1979): mapped from all but extreme N Mo.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Nelson (1977): Reynolds Co. [as Parmelia hypotropa].
Wetmore (1992a): Taney Co. [as Parmelia hypotropa]. See Hale (1957a) citation for P. perforatum, and discussions under Parmelia erecta and Parmelia perlata.
Parmotrema louisianae (Hale) Hale
(1992a): Taney Co. [as Parmelia louisianae].
Parmotrema madagascariaceum (Hue) Hale
Ladd (1991): Ripley, Ste. Genevieve, and Shannon cos.; designated as Rare.
Ladd & Wilhelm (1986): Ste. Genevieve Co.
MDC (1991, 1992): designated as Rare. See note under P. xanthinum.
Parmotrema margaritatum (Hue) Hale
Wetmore (1992a): Taney Co. [as Parmelia margaritata].
Wetmore (1992b): Newton Co. [as Parmelia margaritata].
Parmotrema mellissii - attributed to Missouri only on the basis of the shaded general range map in Hale (1979), which touches extreme southeastern Missouri. Given the level of resolution of these maps, and the lack of confirming specimens, this species is excluded from the flora.
Parmotrema michauxianum (Zahlbr.) Hale
Gier & Kendrick (1972): Barry, Dade, Jasper, and Newton cos., also literature report [as Parmelia michauxiana].
Hale (1969): mapped statewide [as Parmelia michauxiana].
Hale (1979): mapped statewide. Note: the Reynolds Co. specimen cited by Nelson (1977) [as Parmelia michauxiana], Nelson B16 (MOR), is a mixture of Myelochroa galbina and Hypotrachyna livida.
Parmotrema perforatum (Jacq.) A. Massal.
Berry (1937): Franklin, Gasconade, and Washington cos. [as Parmelia perforata].
Berry (1941): Iron, St. Francois, and Wayne cos. [as Parmelia erecta]; Franklin and Wayne cos. [as Parmelia perforata].
C. Culberson (1969): literature report [as Parmelia perforata].
W. Culberson (1973): maps from S1/4 Mo. [as Parmelia perforata].
W. Culberson (1986): maps 4 records from S Mo.
W. Culberson & C. Culberson (1973): maps 5 records in SW Mo. [as Parmelia perforata].
Dey (1976): mapped from SE Mo., based on literature report [as Parmelia perforata].
Dey (1979): mapped from SE Mo., based on literature report.
Gier & Jones (1954): literature report [as Parmelia erecta].
Gier & Kendrick (1972): literature report [as Parmelia perforata].
Hale (1957a): cited from 6 cos. in S1/2 Mo. [as Parmelia erecta]; this includes Parmotrema hypotropum, which is not mentioned in the report; Hale mentions that sorediate and esorediate "phases" were equally abundant. Note that Hale's citation of Parmelia perforata in this report is listed as containing salazinic acid; these reports are referable to Parmotrema eurysacum or possibly Rimelia cetrata.
Hale (1957b): Barry, Douglas, Ozark, Taney, and Texas cos. [as Parmelia hypotropoides].
Hale (1965): 5 records mapped SC Mo. [as Parmelia perforata].
Hale (1969): mapped from S1/2 Mo. [as Parmelia perforata].
Hale (1979): mapped from S1/2 Mo.
Nelson (1977): Reynolds Co. [as Parmelia perforata].
Wetmore (1992a): Taney Co. [as Parmelia perforata]. See Parmelia erecta and Gier & Kendrick (1972) citation of Xanthoparmelia somloënsis.
Parmotrema reticulatum
Rimelia reticulata
Parmotrema subisidiosum
Rimelia subisidiosa
Parmotrema submichauxianum
Parmotrema gardneri
Parmotrema subtinctorium
Rimeliella subtinctoria
Parmotrema tinctorum (Delise ex Nyl.) Hale
Hale (1969): mapped from extreme SE Mo. [as Parmelia tinctorum].
Hale (1979): mapped from extreme SE Mo.
Ladd (1991): Phelps and St. Clair cos.; designated as Endangered.
MDC (1991): designated as Rare.
MDC (1992): designated as Endangered.
Parmotrema ultralucens - attributed to Missouri on the basis of the shaded range map in Hale (1979). Since there are no confirming specimens or corroborating reports, this species is provisionally excluded from the flora. See Parmelia subcrinita.
Parmotrema xanthinum - mapped from southeastern Missouri by Hale (1969, 1979) [the 1969 report as Parmelia xanthina]; the 1969 report forms the basis for the literature report in Gier & Kendrick (1972) [as Parmelia xanthina]. The Gentry Co. report by Gier & Kendrick (1972) is based on Stigers 9226 (MCJ), which is Candelaria concolor. Parmotrema xanthinum lacks gyrophoric acid, which is present in the morphologically analogous Parmotrema madagascariaceum. As pointed out by Almeda & Dey (1973), the presence of gyrophoric acid in some specimens of P. madagascariaceum can best be detected by chromatography, and is not readily revealed by spot tests. All Missouri material examined to date, including several specimens with C medullary reactions, contains gyrophoric acid, and is referable to P. madagascariaceum. In the absence of confirming specimens, Parmotrema xanthinum is provisionally excluded from the Missouri flora.
Patellaria alba - cited from Pike Co. by Pech (1866); it is not known to what taxon this name refers, although Pech lists it as a lichen.
PELTIGERA Willd.
Peltigera sp.
Darigo (1995a): St. Louis Co. See Collema flaccidum.
Peltigera canina (L.) Willd.
AES (1986): Reynolds Co.
Berry (1936): Boone Co.
Berry (1937): Boone, Franklin, Gasconade, Jefferson, and Lincoln cos.
Gier & Jones (1954): literature report [as var. ulorrhiza].
Gier & Kendrick (1972): Jasper and Vernon cos., also literature report; Cole Co. [as var. albescens]; literature report [as var. ulorrhiza].
Hale (1969, 1979): mapped statewide.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Nelson (1911): St. Louis Co.
Nelson (1977): Reynolds Co.
Pech (1866): Pike Co.; this may include material referable to P. rufescens.
Seaward (1978): Salt River basin.
Thomas & Jackson (1985).
Thomson (1950): Greene Co. [as var. ulorrhiza]. See P. degenii.
Peltigera canina var. albescens
Peltigera canina
Peltigera canina var. rufescens f. innovans
Peltigera praetextata
Peltigera canina var. rufescens f. rufescens
Peltigera rufescens
Peltigera canina f. sorediata
Peltigera didactyla
Peltigera canina var. spuria
Peltigera didactyla
Peltigera canina var. spuria f. sorediata
Peltigera didactyla
Peltigera canina var. ulorrhiza
Peltigera canina
Peltigera degenii - cited from McDonald Co. by Gier & Kendrick (1972); the specimen on which this report is based, Palmer M117 (MCJ), is P. canina.
Peltigera didactyla (With.) J. R. Laundon
Gier & Jones (1954): literature reports [as P. canina var. spuria and P. canina var. spuria f. sorediata].
Gier & Kendrick (1972): literature report; literature report [as P. canina f. sorediata].
Hale (1969): mapped from N1/3 Mo. [as P. spuria].
Hale (1979): mapped from NE1/2 Mo. [as P. spuria].
Thomson (1950): Franklin Co. [as P. canina var. spuria f. sorediata]; Sulphur Springs [as P. canina var. spuria] - USGS (1991) lists three Sulphur Springs in Missouri: a town in Jefferson Co. and springs in Miller and Vernon cos. Note: the McDonald Co. specimen cited by Gier & Kendrick (1972) [as P. canina f. sorediata] is based on Lett s.n. 11 October 1970 (MCJ 10549), which is P. rufescens.
Peltigera elisabethae Gyeln.
Ladd et al. (1994): Dade, Oregon, Ripley, Shannon, and Warren cos.
Peltigera evansiana Gyeln.
Gier & Jones (1954): literature report.
Gier & Kendrick (1972): literature report.
Hale (1969): mapped from NE1/5 Mo.
Hale (1979): mapped from E1/3 Mo.
Thomson (1950): Phelps Co.
Peltigera horizontalis - cited from Berry (1936, 1937) from Boone Co., and mapped from extreme SE Mo. by Hale (1979). The 1937 Berry report is the basis for the literature report in Gier & Kendrick (1972). No confirming specimens have been seen, and neither Berry work includes mention of P. polydactylon, which is known to occur in the region. Missouri reports of this species are probably referable to P. polydactylon. The only Gier determination as this taxon that has been located, Gier s.n. 13 September 1974 (MCJ 11697), from Taney Co., is P. rufescens.
Peltigera polydactyla
Peltigera polydactylon
Peltigera polydactyla var. crassoides
Peltigera polydactylon
Peltigera polydactyla f. mucrocarpa
Peltigera polydactylon
Peltigera polydactyla f. typica
Peltigera polydactylon
Peltigera polydactyla var. typica
Peltigera polydactylon
Peltigera polydactyla var. typica f. microcarpa
Peltigera polydactylon
Peltigera polydactylon (Neck.) Hoffm.
Darigo (1995a): Ste. Genevieve Co.
& Jones (1954): literature reports [as P. polydactyla var. crassoides and var. typica f. microcarpa]; Carter Co. [as P. polydactyla var. typica].
Gier & Kendrick (1972): Carter Co., literature report [as P. polydactyla f. typica]; literature report [as P. polydactyla var. crassoides and P. polydactyla f. mucrocarpa].
(1969, 1979): mapped statewide [as P. polydactyla].
Ladd (1996): Phelps Co.
Nelson (1911): St. Louis Co. [as P. polydactyla].
Thomson (1950): St. Francois and Ste. Genevieve cos. [as P. polydactyla var. typica]; Perry Co. [as P. polydactyla var. typica f. microcarpa]; Taney Co. [as P. polydactyla var. crassoides]. See P. horizontalis.
Peltigera praetextata (Flörke ex Sommerf.) Zopf
Gier & Jones (1954): St. Louis Co. [as P. canina var. rufescens f. innovans].
Gier & Kendrick (1972): Livingston and St. Louis cos.
Hale (1969): mapped statewide.
Thomson (1950): Dunklin, Greene, Perry, St. Louis, and Stone cos. [as P. canina var. rufescens f. innovans].
Peltigera rufescens (Weiss) Humb.
Berry (1936, 1937): Boone Co.
Gier & Jones (1954): Camden, Clinton, Jefferson, and St. Louis cos. [as P. canina var. rufescens f. rufescens].
Gier & Kendrick (1972): cited from 6 cos., also literature report; Camden, Clinton, Jefferson, and St. Louis cos., also literature report [as P. canina var. rufescens].
Hus (1908): St. Louis region.
Ladd (1996): Phelps Co.
MBG (1974a, 1974b): Salt River basin.
Nelson (1911): St. Louis Co.
Seaward (1978): Salt River basin.
Thomson (1950): cited from 7 cos. [as P. canina var. rufescens f. rufescens].
Wetmore (1992a): Taney Co. See Pech (1866) citation under P. canina; and notes under P. didactyla and P. horizontalis.
Peltigera spuria
Peltigera didactyla
PELTULA Nyl.
Peltula bolanderi (Tuck.) Wetmore
Wetmore (1992a): Taney Co.
Peltula euploca (Ach.) Poelt
Ladd & Wilhelm (1986): Osage Co. See Collema tunaeforme.
PERTUSARIA DC.
Pertusaria sp.
Seaward (1978): Salt River basin.
Wetmore (1992a): Taney Co. (3 species).
Pertusaria amara (Ach.) Nyl.
Dibben (1980): Barry, Howell, and Pulaski cos.
Ladd (1996): Phelps Co.
Wetmore (1992a): Taney Co.
Pertusaria communis - see discussion under P. pertusa.
Pertusaria hymenea - cited from the Salt River basin by MBG (1974b) and by Seaward (1978). According to Dibben (1980), reports of this taxon from North America east of California are misidentifications.
Pertusaria hypothamnolica Dibben
Dibben (1980): cited from 10 cos.
Ladd (1996): Phelps Co.
Wetmore (1992a): Taney Co. See note under P. multipunctoides.
Pertusaria leioplaca DC.
Berry (1937): Jefferson Co.
Gier & Kendrick (1972): literature report.
Wetmore (1992a): Taney Co. [as P. leucostoma].
Pertusaria leucostoma
Pertusaria leioplaca
Pertusaria macounii (I. M. Lamb) Dibben
Ladd et al. (1994): Camden, Iron, Shannon, Stone, and Wayne cos.
Pertusaria marginata
Pertusaria propinqua
Pertusaria multipuncta - according to Esslinger & Egan (1995), North American reports of this taxon are based on misidentified material. Local reports of P. multipuncta are usually referable to P. multipunctoides.
Pertusaria multipunctoides Dibben
Berry (1937): Boone, Franklin, Gasconade, and Lincoln cos. [as P. multipuncta]. Note: the Berry (1936) report of this species [as P. multipuncta] is based on Berry 304C (UMO), which is P. paratuberculifera. This is the only one of 5 specimens cited by Berry (1937) that has been located. The Camden Co. reports by Gier & Jones (1954) and Gier & Kendrick (1972) [both as P. multipuncta] are based on Jones 13 (MCJ), which is P. hypothamnolica.
Pertusaria neoscotica I. M. Lamb
Dibben (1980): Washington Co.
Ladd (1996): Phelps Co.
Pertusaria ostiolata Dibben
Ladd et al. (1994): cited from 6 cos. S1/3 Mo.
Pertusaria paratuberculifera Dibben
Dibben (1980): cited from 7 cos.
Ladd (1996): Phelps Co.
Wetmore (1992a): Taney Co. See note under P. multipunctoides.
Pertusaria pertusa - cited from Washington Co. by Berry (1937), and from Missouri by Fink (1899) and Tuckerman (1882) [both as P. communis]; Gier & Kendrick (1972) cite the Fink report. According to Esslinger & Egan (1995), North American reports of this taxon are based on misidentifications. Dibben (1980) lists several common species of Pertusaria to which the name P. pertusa has been applied, so translation of literature reports attributing this species to Missouri is not possible. Some early Missouri collections determined as P. communis are referable to P. plittiana.
Pertusaria plitteana [sensu Gier & Kendrick (1972)]
Pertusaria plittiana
Pertusaria plittiana Erichsen
Dibben (1980): Howell, Iron, St. Francois, and Wayne cos.
Gier & Kendrick (1972): Vernon Co. [as P. "plitteana"].
Ladd (1996): Phelps Co. See discussion under P. pertusa.
Pertusaria propinqua Müll. Arg.
Berry (1937): St. Louis Co. [as P. marginata].
Dibben (1980): Phelps, Wayne, and Wright cos.
Gier & Kendrick (1972): literature report [as P. marginata].
Ladd (1996): Phelps Co.
Pertusaria pulchella Malme
Ladd et al. (1994): Carter, Oregon, Ozark, and Shannon cos.
Pertusaria pustulata (Ach.) Duby
Ladd (1996): Phelps Co.
Wetmore (1992a): Taney Co. Note: this species was also cited from Boone Co. by Berry (1936, 1937); the 1937 report forms the basis for the literature report in Gier & Kendrick (1972). The specimen on which these reports are based, Berry 312 (UMO), is Arthothelium spectabile.
Pertusaria subpertusa Brodo
Ladd et al. (1994): Carter and Shannon cos.
Pertusaria tetrathalamia (Fée) Nyl.
Berry (1937): Johnson and St. Louis cos.
Dibben (1980): Howell, Johnson, and St. Louis cos.
Gier & Kendrick (1972): literature report
Ladd (1996): Phelps Co.
Wetmore (1992a): Taney Co.
Pertusaria texana Müll. Arg.
Atienza & Hawksworth (1994): literature report.
Dibben (1980): Barry, Jefferson, and St. Louis cos.
Ladd (1996): Phelps Co.
Wetmore (1992a): Taney Co.
Pertusaria trachythallina Erichsen
Ladd et al. (1994): Shannon Co.
Pertusaria valliculata Dibben
Dibben (1980): Barry Co.
Wetmore (1992a): Taney Co.
Pertusaria velata (Turner) Nyl.
AES (1986): Reynolds Co.
Berry (1937): Franklin Co.
Dibben (1980): cited from 11 cos.
Gier & Kendrick (1972): literature report.
Ladd (1996): Phelps Co.
Wetmore (1992a): Taney Co.
PHAEOCALICIUM A. F. W. Schmidt
Phaeocalicium polyporaeum (Nyl.) Tibell
Ladd et al. (1994): Shannon Co.
Phaeographis eulectra
Graphis eulectra
PHAEOPHYSCIA Moberg
Phaeophyscia adiastola (Essl.) Essl.
Esslinger (1977a): 2 or 3 records mapped from St. Louis area [as Physcia adiastola].
Hale (1979): mapped from SE1/3 Mo.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park. See Phaeophyscia ciliata, Phaeophyscia orbicularis, and Physcia virella.
Phaeophyscia cernohorskyi (Nádv.) Essl.
Esslinger (1978): 5 mapped records from SE2/3 Mo.
Gier & Kendrick (1972): literature report [as Physcia orbicularis f. albociliata].
Hale (1979): mapped statewide.
Ladd (1993a): St. Louis Co.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Thomson (1963): cited from 8 cos. in S Mo. [as Physcia orbicularis f. albociliata].
Wetmore (1992a): Taney Co. See discussion under Physcia virella.
Phaeophyscia ciliata (Hoffm.) Moberg
Berry (1936): Boone Co. [as Physcia obscura]; this may include Phaeophyscia hirtella.
C. Culberson (1969): literature report [as Physcia ciliata].
Dey (1976): mapped statewide, based on literature report [as Physcia ciliata].
Dey (1979): mapped statewide, based on literature report.
Esslinger (1978): Cooper Co.
Fink (1900): Cole Co. [as Physcia obscura]; may include P. hirtella.
Gier & Kendrick (1972): Texas Co., also literature report [as Physcia ciliata].
Hale (1957a): Barry, Crawford, and Ozark cos. [as Physcia ciliata]; possibly including P. hirtella.
Hale (1969): mapped statewide [as Physcia ciliata].
Hale (1979): mapped statewide.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Pech (1866): Pike Co. [as Parmelia ulotrix]; may include P. hirtella.
Thomson (1963): Cooper, Franklin, Jefferson, Ozark, and St. Clair cos. [as Physcia ciliata].
Wetmore (1992a): Taney Co.
Wetmore (1992b): Newton Co. See discussion under Physcia virella. Note: this species was also reported from Dade Co. by Gier & Kendrick (1972) [as Physcia ciliata], but the specimen on which this report is based, Murray 96 (MCJ), is Phaeophyscia adiastola.
Phaeophyscia hirsuta (Mereschk.) Essl.
Wetmore (1992b): Newton Co.
Phaeophyscia hirtella Essl.
AES (1986): Reynolds Co.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Thomson (1963): Cole, Crawford, and Vernon cos. [as Physcia ciliata f. fibrillosa]. See Physcia setosa, and Berry (1936), Fink (1900), Hale (1957a), and Pech (1866) citations under P. ciliata.
Phaeophyscia imbricata (Vain.) Essl.
Esslinger (1978): 2 records mapped from EC Mo.
Gier & Kendrick (1972): literature report [as Physcia lacinulata and Physcia ciliata f. squamulosa].
Hale (1969): mapped from SE1/2 Mo. [as Physcia lacinulata].
Hale (1979): mapped from E1/3 Mo.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Thomson (1963): Boone and Washington cos. [as Physcia ciliata f. squamulosa].
Wetmore (1992a): Taney Co.
Phaeophyscia insignis (Mereschk.) Moberg
Esslinger (1985): Christian Co. See Phaeophyscia orbicularis.
Phaeophyscia orbicularis (Neck.) Moberg
Gier & Jones (1954): Saline Co. [as Physcia orbicularis].
Gier & Kendrick (1972): Laclede, McDonald, and Saline cos., also literature report [as Physcia orbicularis].
Hale (1957a): cited from 11 cos. in S1/2 Mo. [as Physcia orbicularis].
Hale (1969): mapped statewide [as Physcia orbicularis].
Ladd (1996): Phelps Co.
MBG (1974b): Salt River basin [as Physcia orbicularis].
Nelson (1977): Reynolds Co. [as Physcia orbicularis].
Seaward (1978): Salt River basin [as Physcia orbicularis].
Thomson (1963): cited from 12 cos. [as Physcia orbicularis f. orbicularis].
Wetmore (1992a): Taney Co. Note: all of these reports may include or consist entirely of material referable to Phaeophyscia adiastola, P. insignis, P. pusilloides, P. rubropulchra, and/or Physciella chloantha, and possibly Phaeophyscia cernohorskyi. Phaeophyscia orbicularis sensu stricto is known to occur in Missouri, but is rare and usually saxicolous, whereas most of the above citations refer to corticolous populations. This species is also cited from Barry and Newton cos. by Gier & Kendrick (1972) [as Physcia orbicularis], but the Barry Co. report is based on Sauer s.n. 11 May 1963 (MCJ 10664), which is Physconia detersa, and the Newton Co. report is based on Kendrick s.n. 27 February 1971 (MCJ 10954), which is P. cernohorskyi.
Phaeophyscia pusilloides (Zahlbr.) Essl.
Esslinger (1978): 4 records mapped from EC and SW Mo.
Hale (1979): mapped statewide.
Ladd (1993a): St. Louis Co.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Wetmore (1992a): Taney Co. See discussions under Phaeophyscia orbicularis, Physcia americana, and Physcia virella.
Phaeophyscia rubropulchra (Degel.) Essl.
Berry (1937): Boone and Franklin cos. [as Physcia endochrysea].
Darigo (1995a): St. Louis Co.
Gier & Kendrick (1972): literature report [as Physcia orbicularis f. rubropulchra].
Hale (1979): mapped from S3/4 Mo.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Thomson (1963): Iron and Newton cos. [as Physcia orbicularis f. rubropulchra].
Wetmore (1992a): Taney Co. See discussions under Phaeophyscia orbicularis, Physcia millegrana, Pyxine sorediata, and Pyxine subcinerea.
PHLYCTIS Wallr.
Phlyctis argena (Spreng.) Flot.
Ladd et al. (1994): Reynolds and Wayne cos.
PHYSCIA (Schreb.) Michx.
Physcia adglutinata
Hyperphyscia adglutinata
Physcia adiastola
Phaeophyscia adiastola
Physcia adscendens (Fr.) H. Olivier
Ladd et al. (1994): Grundy, Livingston, and Ste. Genevieve cos.
Physcia aipolia (Ehrh. ex Humb.) Fürnr.
C. Culberson (1969): literature report.
Gier & Jones (1954): Saline Co.; Clay Co. [as f. acrita].
Gier & Kendrick (1972): Andrew, Clay, and Saline cos., also literature report; Clay Co., also literature report [as var. acrita].
Hale (1957a): Ozark and Taney cos.
Hale (1969, 1979): mapped statewide.
Thomson (1963): Clay, Johnson, Ozark, and Vernon cos.; Taney Co. [as f. cercida].
Wetmore (1992a): Taney Co.
Wetmore (1992b): Newton Co. Note: there is a possibility that some of these reports may include or consist of material referable to P. pumilior; see discussion in Harris (1990). See P. astroidea, P. pulverulenta, and P. stellaris entries.
Physcia aipolia f. acrita
Physcia aipolia
Physcia aipolia var. acrita
Physcia aipolia
Physcia aipolia f. cercida
Physcia aipolia
Physcia alba - according to Esslinger & Egan (1995), this species does not occur in North America. Local reports are referable to Physcia pumilior.
Physcia americana G. Merr.
Berry (1936): Boone Co. [as P. astroidea and P. caesia].
Berry (1937): Boone and Johnson cos. [as P. astroidea and P. caesia]; Jefferson, St. Louis, and Washington cos. [as P. astroidea].
C. Culberson (1969): literature report [as P. tribacoides].
Gier & Kendrick (1972): Laclede, Newton, and Vernon cos., also literature report [as P. tribacoides].
Hale (1957a): cited from 12 cos. in S1/2 Mo [as P. tribacoides].
Hale (1969): mapped statewide [as P. tribacoides].
Hale (1979): mapped statewide.
Hawksworth & Booth (1976): Salt River basin [as P. tribacoides].
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Seaward (1978): Salt River basin [as P. tribacoides].
Thomson (1963): cited from 16 cos. [as P. tribacoides].
Wetmore (1992a): Taney Co.
(1992b): Newton Co. Note: the Reynolds Co. specimen cited by Nelson (1977) is Punctelia perreticulata. Some of Berry's reports of Physcia caesia may be referable to Physconia detersa; see discussion under Physcia caesia. The Cedar Co. report in Gier & Kendrick (1972) [as P. tribacoides] is based on Kendrick s.n. 29 March 1971 (MCJ 11027), which is a mixture of Phaeophyscia pusilloides and Physciella chloantha.
Physcia astroidea [
P. clementei (Sm.) Lynge] - local reports are referable to P. americana. A Berry determination as this taxon, Berry 158A (UMO), is P. aipolia, and another Berry determination, Berry 2036 (UMO) is P. americana. Both of these specimens are from Boone Co.
Physcia caesia - this species is cited from Boone Co. by Berry (1936) and from Boone and Johnson cos. by Berry (1937). Neither of these publications includes reference to the morphologically similar P. americana, which is common throughout Missouri. Both Berry reports are referable to P. americana. Although some Berry determinations as this species are P. americana, a Johnson Co. specimen collected as P. caesia, Berry 615 (US), is Physconia detersa. According to the map in Hale (1979), Physcia caesia is a northern and western species not occurring within 250 miles of Missouri. Thomson (1963) maps P. caesia from New England, the Great Lakes north of Lake Michigan, and the western United States, with one record in central Kansas.
Physcia chloantha
Physciella chloantha
Physcia ciliata
Phaeophyscia ciliata
Physcia ciliata f. fibrillosa
Phaeophyscia hirtella
Physcia ciliata f. squamulosa
Phaeophyscia imbricata
Physcia detersa
Physconia detersa
Physcia dubia (Hoffm.) Lettau
Wetmore (1992a): Taney Co.
Wetmore (1992b): Newton Co. Note: this species was also cited from Franklin and Gasconade cos. by Berry (1937), Iron Co. by Fink (1919), and Barry Co. by Gier & Jones (1954) and Gier & Kendrick (1972) [all as P. teretiuscula]. All Missouri reports of P. dubia except those of Wetmore (1992a, 1992b) are probably based on misidentifications. To date, all Missouri specimens labelled as P. teretiuscula that I have seen are Speerschneidera euploca. I have not examined the Wetmore specimens. Local reports may also be referable to Physciella chloantha or Physcia subtilis. According to the map in Hale (1979), the closest station to Missouri for P. dubia is in the Upper Peninsula of Michigan.
Physcia endochrysea
Phaeophyscia rubropulchra
Physcia frostii
Dirinaria frostii
Physcia grisea
Physconia detersa
Physcia grisea f. farrea
Physconia detersa
Physcia halei J. W. Thomson
Gier & Kendrick (1972): McDonald Co., also literature report.
Hale (1969): mapped from SE1/2 Mo.
Hale (1979): mapped from SE 3/5 Mo.
Ladd & Nelson (1982).
Ladd & Wilhelm (1983): Bennett Spring State Park.
Nelson (1977): Reynolds Co.
Thomson (1963): Crawford Co.
Physcia hispida [
P. tenella (Scop.) DC.] - cited from Franklin Co. by Berry (1937), presumably based upon a misidentification. Thomson (1963) cited specimens from no closer to Missouri than North Dakota and northern portions of Minnesota and Michigan.
Physcia lacinulata
Phaeophyscia imbricata
Physcia lithotodes [
Phaeophyscia endococcina (Körb.) Moberg] cited from Franklin Co. by Berry (1937), but no specimens have been located. Thomson (1963, 1984) restricts the range of this species to areas far to the north and west of Missouri.
Physcia luganensis
Physciella chloantha
Physcia melanchra
Physciella melanchra
Physcia millegrana Degel.
AES (1986): Reynolds Co.
Berry (1936): Boone Co. [as P. tribacia].
Berry (1937): Boone, Franklin, and St. Louis cos. [as P. tribacia].
C. Culberson (1969): literature report.
Darigo (1995a): St. Louis Co.
Gier & Kendrick (1972): Cass Co., also literature report.
Hale (1957a): Barry and Crawford cos.
Hale (1969): mapped statewide except central Ozarks.
Hale (1979): mapped from SE border.
Ladd (1993a): St. Louis Co.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
MBG (1974b): Salt River basin.
Nelson (1977): Reynolds Co.
Seaward (1978): Salt River basin.
Thomson (1963): cited from 9 cos.
Wetmore (1992a): Taney Co.
Wetmore (1992b): Newton Co. Note: this species was also cited from Newton Co. by Gier & Kendrick (1972), but the specimen on which this report is based, Kendrick s.n. 27 February 1971 (MCJ 10953), is Phaeophyscia rubropulchra.
Physcia obscura
Phaeophyscia ciliata
Physcia orbicularis
Phaeophyscia orbicularis; but see discussion under Phaeophyscia orbicularis.
Physcia orbicularis f. albociliata
Phaeophyscia cernohorskyi
Physcia orbicularis f. orbicularis - see Physcia orbicularis
Physcia orbicularis f. rubropulchra
Phaeophyscia rubropulchra
Physcia phaea - the only basis for attributing this lichen to Missouri is the imprecise, shaded range map in Hale (1979). Wilhelm & Ladd (1985) cite a specimen from Jackson Co., Illinois, which borders Missouri, so this taxon may someday be discovered in the state.
Physcia pulverulenta [
Physconia distorta (With.) J. R. Laundon ] - Esslinger & Egan (1995) note that North American reports of this taxon are base on misidentifications. Hale (1979) tentatively refers reports of this species [as Physconia pulverulenta] from the eastern United States to Anaptychia palmulata [his A. "palmatula"], but all Missouri material previously identified as Physcia pulverulenta is referable to Physconia detersa, except for a Jasper Co. specimen cited by Gier & Kendrick, Kendrick s.n. 20 November 1970 (MCJ 10990), which is Physcia aipolia.
Physcia pumilior R. C. Harris
Harris (1990). Note: the Cedar Co. report in Gier & Kendrick (1972) [as P. alba] is based on Kendrick s.n. 29 March 1971 (MCJ 11029), which is Hyperphyscia syncolla. The Newton Co. report in Gier & Kendrick (1972) [as P. alba] is based on Kendrick s.n. 21 March 1970 (MCJ 11033), which is Hyperphyscia adglutinata. See notes under P. aipolia and P. stellaris.
Physcia setosa [
Phaeophyscia hispidula (Ach.) Essl.] - cited from Newton Co. by Gier & Kendrick (1972). This report is based on Kendrick s.n. 24 November 1970 (MCJ 10991), which is Phaeophyscia hirtella.
Physcia stellaris (L.) Nyl.
Berry (1936): Boone Co.
Berry (1937): cited from 8 cos.
Brodo (1968): mapped statewide.
C. Culberson (1969): literature report.
Gier & Kendrick (1972): Boone, Cass, and Dade cos., also literature report.
Hale (1957a): cited from 7 cos. in S1/2 Mo.
Hale (1969, 1979): mapped statewide.
Hus (1908): St. Louis region.
Ladd (1993a): St. Louis Co.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Nelson (1911): Jefferson Co.
Nelson (1977): Reynolds Co.
Seaward (1978): Salt River basin.
Thomas & Jackson (1985).
Thomson (1963): cited from 16 cos.
Wetmore (1992a): Taney Co. Note: some of these reports may include or consist of material referable to P. pumilior; see discussion in Harris (1990). The Vernon Co. report in Gier & Kendrick (1972) is based on Kendrick s.n. 20 February 1971 (MCJ 10957), which is Physcia aipolia. See Parmelia physodes.
Physcia subtilis Degel.
Gier & Kendrick (1972): Jasper and Newton cos., also literature report.
Hale (1969): mapped statewide.
Hale (1979): mapped from SE Mo.
Ladd (1996): Phelps Co.
Ladd & Nelson (1982).
Ladd & Wilhelm (1983): Bennett Spring State Park.
Thomson (1963): Franklin and Iron cos.
Wetmore (1992a): Taney Co. Note: some local reports of P. dubia may be referable here; see discussion under that species.
Physcia syncolla
Hyperphyscia syncolla
Physcia teretiuscula - see Physcia dubia
Physcia tribacia - local reports are referable to Physcia millegrana.
Physcia tribacoides
Physcia americana
Physcia virella [
Phaeophyscia adiastola] - cited from Boone Co. by Berry (1936), and from Boone, Franklin, and Jefferson cos. by Berry (1937). These collections are all described as corticolous, and may include or consist of Phaeophyscia pusilloides, which is usually corticolous, as opposed to Phaeophyscia adiastola, which is prevailingly saxicolous with us. Some Berry collections determined as this species are Phaeophyscia cernohorskyi and Phaeophyscia ciliata.
PHYSCIELLA Essl.
Physciella chloantha (Ach.) Essl.
Esslinger (1978): one record mapped from SW Mo. [as Physcia luganensis].
Esslinger (1986): Christian Co.
Hale (1979): mapped statewide [as Physcia chloantha].
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park [as Physcia chloantha].
Wetmore (1992a): Taney Co. [as Physcia chloantha]. Note: some local reports of Physcia dubia may be referable here. See discussions under Phaeophyscia orbicularis, Physcia americana, and Pyxine sorediata.
Physciella melanchra (Hue) Essl.
Ladd (1996): Phelps Co.
Wetmore (1992a): Taney Co. [as Physcia melanchra].
Physciopsis adglutinata
Hyperphyscia adglutinata
Physciopsis syncolla
Hyperphyscia syncolla
PHYSCONIA Poelt
Physconia detersa (Nyl.) Poelt
Berry (1936, 1937): Boone Co. [as Physcia pulverulenta].
C. Culberson (1969): literature report [as Physcia grisea].
Gier & Jones (1954): Camden and Clay cos. [as Physcia grisea].
Gier & Kendrick (1972): Camden and Clay cos., also literature report [as Physcia grisea]; literature report [as Physcia pulverulenta].
Hale (1957a): cited from 11 cos. in S1/2 Mo. [as Physcia grisea].
Hale (1969): mapped statewide [as Physcia grisea].
Hale (1979): mapped statewide.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
MBG (1974a, 1974b): Salt River basin [as Physcia detersa]; erroneously cited as new to Missouri.
Seaward (1978): Salt River basin.
Thomson (1963): cited from 22 cos. [as Physcia grisea]; Stone Co. [as Physcia grisea f. farrea].
Wetmore (1992a): Taney Co.
Wetmore (1992b): Newton Co. Note: most of these reports may include Physconia kurokawae; see discussion in Ladd et al. (1994). See discussions under Phaeophyscia orbicularis, Physcia caesia, and Physcia pulverulenta.
Physconia kurokawae Kashiw.
Ladd et al. (1994): cited from 13 cos. statewide. See discussion under P. detersa.
Physma luridum
Pannaria lurida
Physolinum monile - see Coenogonium missouriense
Placodium albescens - cited from Pike Co. by Pech (1866); it is not known to what taxon this name refers.
Placodium aurantiacum
Caloplaca flavorubescens
Placodium citrinum
Caloplaca citrina
Placodium ferrugineum
Caloplaca ferruginea
Placodium galactophyllum
Caloplaca galactophylla
Placodium vitellinum var. aurellum - see Candelariella aurella
PLACYNTHIELLA Elenkin
Placynthiella icmalea (Ach.) Coppins & P. James
Wetmore (1992a): Taney Co.
PLACYNTHIUM (Ach.) Gray
Placynthium nigrum (Huds.) Gray
Gier & Kendrick (1972): literature report.
Henssen (1963): cited from 6 cos.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Nelson (1985).
Wetmore (1992a): Taney Co. See comments under P. stenophyllum.
Placynthium petersii (Nyl.) Burnham
Berry (1937): Jefferson Co. [as Pterygium petersii].
Gier & Kendrick (1972): literature report.
Placynthium stenophylla [sensu Gier & Jones (1954)] - see Placynthium stenophyllum
Placynthium stenophyllum (Tuck.) Fink - reported from Saline Co. by Gier & Jones (1954) [as P. "stenophylla"] and Gier & Kendrick (1972). The specimen on which these reports are based, Jones 18 (MCJ), is P. nigrum.
Polyblastiopsis fallax - see Arthopyrenia lapponina
Polychidium umhausense [
Dendriscocaulon umhausense (Auersw.) Degel.] - local reports are referable to Dendriscocaulon intricatulum; see James & Henssen (1976).
POLYSPORINA Vezda
Polysporina simplex (Davies) Vezda
Fink (1919): Iron Co. [as Biatorella simplex].
Gier & Kendrick (1972): literature report mistakenly attributed to Fink (1900) [as Sarcogyne simplex].
PORINA Müll. Arg.
Porina cestrensis (Tuck. ex E. Michener) Müll. Arg.
Ladd et al. (1994): cited from 8 cos SE1/2 Mo. Note: Harris (1995b) proposes the name Trichothelium cestrense (E. Michener) R. C. Harris for this taxon.
PORPIDIA Körb.
Porpidia albocaerulescens (Wulfen) Hertel & Knoph
Fink (1935) [as Lecidea albocaerulescens].
Gier & Jones (1954): literature report [as Lecidea albocaerulescens].
Gier & Kendrick (1972): literature report mistakenly attributed to Berry (1937) [as Lecidea albocaerulescens].
Gowan (1989): 2 records mapped from SW and EC Mo. (stictic acid type; esorediate).
Wetmore (1992a): Taney Co. Note: this taxon was also cited from Boone Co. by Berry (1936) [as Lecidea albocaerulescens], but this report must be based on a misidentification, since he cites the substrate as corticolous, whereas all populations seen in Missouri are saxicolous.
Porpidia crustulata (Ach.) Hertel & Knoph
Gowan (1989): one record mapped in extreme S Mo.
Porpidia tahawasiana Gowan
Ladd (1996): Phelps Co.
Ladd et al. (1994): Reynolds and Ste. Genevieve cos.
PRONECTRIA Clem.
Pronectria oligospora Lowen & Rogerson*
Lowen (1995): Ste. Genevieve Co.
PROTOBLASTENIA (Zahlbr.) J. Steiner
Protoblastenia rupestris (Scop.) J. Steiner
Ladd (1996): Phelps Co.
Wetmore (1992a): Taney Co.
PSEUDOCYPHELLARIA Vain.
Pseudocyphellaria aurata (Ach.) Vain.
Gier & Kendrick (1972): mention "should" occur in Mo.
Hale (1969): mapped from SE1/3 Mo.
Hale (1979): mapped from extreme SE Mo.
Ladd (1991): Dunklin and Oregon cos.; designated as Endangered.
MDC (1991, 1992): designated as Endangered.
Pseudoparmelia sp. [sensu Morgan (1987)]
Flavoparmelia sp.
Pseudoparmelia baltimorensis
Flavoparmelia baltimorensis
Pseudoparmelia caperata
Flavoparmelia caperata
Pseudoparmelia caroliniana
Canoparmelia caroliniana
Pseudoparmelia crozalsiana
Canoparmelia crozalsiana
Pseudoparmelia texana
Canoparmelia texana
PSORA Hoffm.
Psora decipiens (Hedw.) Hoffm.
Fink (1899) [as Biatora decipiens].
Gier & Jones (1954): St. Louis Co. [as Lecidea decipiens].
Gier & Kendrick (1972): St. Louis Co., also literature report [as Lecidea decipiens].
Hale (1979): mapped statewide.
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1983): Bennett Spring State Park.
Tuckerman (1888) [as Biatora decipiens].
Psora icterica (Mont.) Müll. Arg.
Ladd (1991): Cedar and Lawrence cos.; designated as Endangered.
Ladd & Wilhelm (1986): Cedar and Lawrence cos.
MDC (1991, 1992): designated as Endangered.
Nelson (1985).
Psora pseudorussellii Timdal
Gier & Kendrick (1972): literature report [as Lecidea rubiformis].
Hale (1969): mapped statewide [as Lecidea rubiformis]; may also include Psora russellii.
Ladd (1996): Phelps Co.
Ladd & Nelson (1982) [as P. russellii].
Ladd & Wilhelm (1983): Bennett Spring State Park [as P. russellii].
Morgan (1987) [as P. russellii].
Nelson (1977): Reynolds Co. [as Lecidea rubiformis].
Nelson (1985) [as P. russellii].
Timdal (1986): mapped from SW and EC Mo.
Tuckerman (1888) [as Biatora russellii].
Wetmore (1992a): Taney Co. Note: as discussed by Timdal (1986); P. pseudorussellii is saxicolous as opposed to the almost invariably terricolous P. russellii sensu stricto. Previous reports of P. russellii that are described as wholly or partially saxicolous are referred here. Field experience in Missouri indicates that by far the more common of the two taxa is P. pseudorussellii. See Lecidea rubiformis.
Psora russellii (Tuck.) A. Schneid.
Fink (1899) [as Biatora russellii]; this may include or consist entirely of Psora pseudorussellii.
Hale (1979): mapped from NW and SE Mo.
Nelson (1911): "Mineke" [as Biatora russellii]; this report probably includes or consists entirely of P. pseudorussellii.
Timdal (1986): one record mapped from SW Mo.
Wetmore (1992a): Taney Co. Note: reports of this taxon by Ladd & Nelson (1982), Ladd & Wilhelm (1983), Morgan (1987), Nelson (1985), and Tuckerman (1888) are referred to P. pseudorussellii, which see. See discussion under Lecidea russula and L. rubiformis.
Psora tuckermanii - local reports are referable to Lecidea lurida.
PSOROTICHIA A. Massal.
Psorotichia schaereri (A. Massal.) Arnold
Ladd et al. (1994): Buchanan, Dade, Greene, and Lewis cos.
PSORULA Gotth. Schneid.
Psorula rufonigra (Tuck.) Gotth. Schneid.
Fink (1899) [as Biatora rufonigra].
Gier & Kendrick (1972): literature report [as Lecidea rufonigra].
Ladd (1996): Phelps Co.
Ladd & Wilhelm (1986): cited from 10 cos.
Tuckerman (1888) [as Biatora rufonigra].
Wetmore (1992a): Taney Co.
Pterygium petersii
Placynthium petersii
PUNCTELIA Krog
Punctelia bolliana (Müll. Arg.) Krog
Berry (1941): Wayne Co. [as Parmelia bolliana].
W. Culberson (1962): mapped statewide [as Parmelia frondifera].
W. Culberson & C. Culberson (1956): mapped from St. Louis area and SW Mo. [as Parmelia bolliana strain 4].
Gier & Jones (1954): literature report [as Parmelia bolliana].
Gier & Kendrick (1972): Camden, Jasper, Laclede, and Vernon cos., also literature report [as Parmelia bolliana].
Hale (1957a): Ozark and Stone cos. [as Parmelia bolliana].
Hale (1969): mapped statewide [as Parmelia bolliana].
Hale (1979): mapped statewide [as Parmelia bolliana].
Wetmore (1992a): Taney Co. [as Parmelia bolliana].
Wetmore (1992b): Newton Co. [as Parmelia bolliana]. Note: many of the pre-1969 citations listed above may include or refer entirely to Punctelia semansiana. The Reynolds Co. specimen cited by Nelson (1977)[as Parmelia bolliana] and the Boone Co. specimen cited by Berry (1936, 1937) [as Parmelia frondifera], Drouet 396D (UMO), are Hypotrachyna livida. See discussion of Seaward (1978) citation under Punctelia semansiana, and discussion under Parmelia borreri.
Punctelia missouriensis G. Wilh. & Ladd
Ladd (1996): Phelps